（1）根际理论创新与实践应用：以根际为核心，探究多变环境下植物根系-土壤-微生物互作过程、调控机制及其生态学意义；精准调控植物根系-土壤-微生物多功能体互惠协作，实现农林特定生态功能目标。

（2）退化生态系统恢复与保护学：以西南山地退化生态系统为对象，基于生态系统种间关系、物质循环等生态过程的基础研究，研发攻关退化生态系统结构优化和功能提升的技术体系。

2025年

lHan Yang, Peipei Zhang*, Guangru Wang, Jipeng Wang, Dungang Wang, Min Li, Huajun Yin*. Root exudate chemodiversity bridges acquisitive-conservative strategy synergy between roots and rhizosphere microbes in a subtropical forest. Ecology Letters, 2025

lPeipei Zhang, Junxiang Ding, Deliang Kong, Huajun Yin*. Rhizospheric traits and plant functioning belowground. Plant Communications, 2025, 101579

lMin Li, Jipeng Wang*, Na Li, Qitong Wang, Peipei Zhang, Qiuhong Feng, Huajun Yin*. The rhizosphere contributes disproportionately to free-living nitrogen fixation in subalpine forest soils. Soil Biology and Biochemistry, 2025, 200: 109641

lHan Yang, Peipei Zhang*, Qitong Wang, Shaojun Deng, Guangru Wang, Xinjun Zhang, Ruihong Wang, Huajun Yin*. Shifts in root exudate composition coordinate with root resource conservation along an elevation gradient. Journal of Ecology, 2025, 113: 1970–1984

lDeyi Wang, Hans Jacquemyn, Guangru Wang, Yongping Kou, Junxiang Ding, Peipei Zhang, Huajun Yin*. Evolutionary history and environmental filtering jointly structure ectomycorrhizal fungal communities across Qinghai-Tibetan Pinaceae forests. Journal of Plant Ecology, 2025, rtaf159

lDeyi Wang*, Vincent S. F. T. Merckx, Hans Jacquemyn, Sofia I. F. Gomes. Mycorrhizal communities in Orchidaceae are likely shaped by plant trophic mode and biogeography but not phylogeny. Plant Diversity, 2025

lXiaohu Wang, Yutian Wu, Wenqiang Zhao, Huajun Yin*. The rhizosphere and hyphosphere differentially regulate microbiomes and nutrient cycling within soil aggregates in an ectomycorrhiza-dominated forest. Geoderma, 2025, 463: 117570

lChenguang Gao, T. Martijn Bezemer, Jia Liu, Deyi Wang, Jipeng Wang, Shaojun Deng, Dungang Wang, Huajun Yin*. Distinct roles of bacteria and fungi in driving rhizosphere and bulk soil multifunctionality of Abies georgei in an alpine forest. BMC Plant Biology, 2025, 25: 1602

lWentong Gao, Qitong Wang*, Na Li, Ruihong Wang, Xinjun Zhang, Huajun Yin*. Nitrogen addition enhances stable soil carbon accumulation during ectomycorrhizal hyphae decomposition. Plant and Soil, 2025, 511: 521-534

lDungang Wang, Shaojun Deng, Han Yang, Na Li, Qiuhong Feng, Jia Liu*, Huajun Yin. The microbial network exhibits higher complexity in the rhizosphere than in bulk soils along elevational gradients in the alpine forests. Applied Soil Ecology, 2025, 213: 106264

lGuangru Wang, Han Yang, Peipei Zhang*, Qitong Wang, Dungang Wang, Qiuhong Feng, Huajun Yin*. Shifts in ectomycorrhizal exploration types complement root traits in nutrient foraging of alpine coniferous forests along an elevation gradient. Plant and Soil, 2025, 515: 901-914

lHan Yang, Peipei Zhang*, Guangru Wang, Qitong Wang, Dungang Wang, Ruihong Wang, Xinjun Zhang, Huajun Yin*. The synergistic strategy of leaf nitrogen conservation and root nitrogen acquisition in an alpine coniferous forest along an elevation gradient. Plant and Soil, 2025, 514: 2067-2082

2024年

lPeipei Zhang, Junxiang Ding, Qitong Wang, Nate G. McDowell, Deliang Kong, Huajun Yin*. Contrasting coordination of non-structural carbohydrates with leaf and root economic strategies of alpine coniferous forests. New Phytologist, 2024, 243: 580-590. DOI: 10.1111/nph.19678

lShaojun Deng, Qitong Wang*, Dungang Wang, HanYang, Peipei Zhang, Na Li, Xinjun Zhang, Ruihong Wang, Huajun Yin*. The accumulation capacity of microbial residues in the rhizosphere increased along an elevation gradient. Catena, 2024, 238: 107891

lJunxiang Ding, Qitong Wang, Wenjing Ge, Qing Liu, Deliang Kong*, Huajun Yin*. Coordination of leaf and root economic space in alpine coniferous forests on the Tibetan Plateau. Plant and Soil, 2024, 496: 555-568

lWentong Gao, Xiaomin Zhu, Peipei Zhang, Qitong Wang*, Xinjun Zhang*, Huajun Yin. Response of decomposition dynamics of ectomycorrhizal fungal fruit bodies to N addition depends on decomposition stage in a coniferous forest. Applied Soil Ecology, 2024, 193: 105125

lYongping Kou, Junxiang Ding, Huajun Yin*. Temperature governs the community assembly of root-associated ectomycorrhizal fungi in alpine forests on the Qinghai-Tibetan Plateau. Science of The Total Environment, 2024, 176820

lMin Li, Xi He, Peipei Zhang, Ruihong Wang, Jipeng Wang*, Xinjun Zhang*, Huajun Yin. Close linkage between available and microbial biomass phosphorus in the rhizosphere of alpine coniferous forests along an altitudinal gradient. Rhizosphere, 2024, 30: 100904

lDungang Wang, Shaojun Deng, Jipeng Wang, Qiuhong Feng, Jia Liu*, Huajun Yin*. Soil micro-food web complexity drives soil multifunctionality along an elevation gradient. CATENA, 2024, 246: 108464

lDungang Wang, Wentong Gao, Jipeng Wang*, Huajun Yin*. Minimal impact of nitrogen addition on bacterial and fungal communities during fungal necromass decomposition in a subalpine coniferous plantation. Soil Biology and Biochemistry, 2024, 196: 109499

lJipeng Wang, Min Li, Qitong Wang, Ziliang Zhang, Dungang Wang, Peipei Zhang, Na Li, Yiqiu Zhong, Huajun Yin*. Full-factorial resource amendment experiments reveal carbon limitation of rhizosphere microbes in alpine coniferous forests. Biology and Fertility of Soils, 2024, 60: 1181–1186

lQitong Wang, Junxiang Ding, Ziliang Zhang, Chao Liang, Hans Lambers, Biao Zhu, Dungang Wang, Jipeng Wang, Peipei Zhang, Na Li, Huajun Yin*. Rhizosphere as a hotspot for microbial necromass deposition into the soil carbon pool. Journal of Ecology, 2024, 11: 168–179

lQitong Wang, Jipeng Wang, Ziliang Zhang, Min Li, Dungang Wang, Peipei Zhang, Na Li, Huajun Yin*. Microbial metabolic traits drive the differential contribution of microbial necromass to soil organic carbon between the rhizosphere of absorptive roots and transport roots. Soil Biology and Biochemistry, 2024, 197: 109529

lQian Zhao, Jipeng Wang*, Qingqing He, Huajun Yin. Carbon type and quantity regulate soil free-living nitrogen fixation through restructuring diazotrophic community. Applied Soil Ecology, 2024, 202: 105586

2023年

lXiaomin Zhu, Hans Lambers, Wanji Guo, Ziliang Zhang, Huajun Yin*. 2023. Extraradical hyphae exhibit more plastic nutrient-acquisition strategies than roots under nitrogen enrichment in ectomycorrhiza-dominated forests. Global Change Biology, 239: 1651-1664.

lZiliang Zhang, Wanji Guo, Jipeng Wang, Hans Lambers and Huajun Yin*. Extraradical hyphae alleviate nitrogen deposition-induced phosphorus deficiency in ectomycorrhiza-dominated forests. New Phytologist, 2023, 239: 1651–1664.

lJunxiang Ding, Huajun Yin*, Deliang Kong*, Qing Liu, Ziliang Zhang, Qitong Wang, Wanji Guo, Oscar J. Valverde-Barrantes, Junjian Wang, Zhanfeng Liu. Precipitation, rather than temperature drives coordination of multidimensional root traits with ectomycorrhizal fungi in alpine coniferous forests. Journal of Ecology, 2023, DOI: 10.1111/1365-2745.14151

lJunxiang Ding, Wenjing Ge, Qing Liu, Qitong Wang, Deliang Kong*, Huajun Yin*. Temperature drives the coordination between above-ground nutrient conservation and below-ground nutrient acquisition in alpine coniferous forests. Functional Ecology, 2023, 37: 1674–1687

lJunxiang Ding, Qitong Wang, Wenjing Ge, Qing Liu, Deliang Kong*, Huajun Yin*. Coordination of leaf and root economic space in alpine coniferous forests on the Tibetan Plateau. Plant and Soil， 2023

lHan Yang, Peipei Zhang*, Qitong Wang, Shaojun Deng, Xi He, Xinjun Zhang, Ruihong Wang, Qiuhong Feng, Huajun Yin*. Temperature rather than N availability determines root exudation of alpine coniferous forests on the eastern Tibetan Plateau along elevation gradients. Tree Physiology , 2023, 43, 1479-1492

lQitong Wang, Ying Zhang, Peipei Zhang, Na Li, Ruihong Wang, Xinjun Zhang*, Huajun Yin*. Nitrogen deposition induces a greater soil C sequestration in the rhizosphere than bulk soil in an alpine forest. Science of the Total Environment, 2023, 875: 162701

lJunxiang Ding, Qitong Wang, Wenjing Ge, Qing Liu, Deliang Kong, Huajun Yin*. 2023. Coordination of leaf and root economic space in alpine coniferous forests on the Tibetan Plateau. Plant and Soil, 496: 555-568.

2022年

lXiaomin Zhu, Ziliang Zhang, Qitong Wang, Josep Peñuelas, Jordi Sardans, Hans Lambers, Na Li, Qing Liu, Huajun Yin*, Zhanfeng Liu. More soil organic carbon is sequestered through the mycelium pathway than through the root pathway under nitrogen enrichment in an alpine forest. Global Change Biology, 2022, 28, (16), 4947-4961

lQitong Wang, Ziliang Zhang, Xiaomin Zhu, Zhanfeng Liu, Na Li, Juan Xiao, Qing Liu, Huajun Yin*. Absorptive roots drive a larger microbial carbon pump efficacy than transport roots in alpine coniferous forests. Journal of Ecology, 2022, 110, (7), 1646-1655

lPeipei Zhang, Mingzhen Yin, Xinjun Zhang, Qitong Wang, Ruihong Wang*, Huajun Yin*. Differential aboveground-belowground adaptive strategies to alleviate N addition-induced P deficiency in two alpine coniferous forests. Science of the Total Environment, 849 (2022) 157906

2021年

lWentong Gao, Qitong Wang, Xiaomin Zhu, Zhanfeng Liu, Na Li, Juan Xiao, Xiaoping Sun , Huajun Yin*. The vertical distribution pattern of microbial- and plant-derived carbon in the rhizosphere in alpine coniferous forests. Rhizosphere, 2021, 100436

lQitong Wang, Ziliang Zhang, Wanji Guo, Xiaomin Zhu, Juan Xiao, Qing Liu, Huajun Yin*. Absorptive and transport roots differ in terms of their impacts on rhizosphere soil carbon storage and stability in alpine forests. Soil Biology and Biochemistry, 2021, 161, 108379

lWanji Guo, Junxiang Ding, Qitong Wang, Mingzhen Yin, Xiaomin Zhu, Qing Liu, Ziliang Zhang*, Huajun Yin*. Soil fertility controls ectomyc ts in alpine forests receiving nitrogen deposition. Soil Biology and Biochemistry, 2021, 161, 108386

lQitong Wang, Juan Xiao, Junxiang Ding, Tingting Zou, Ziliang Zhang, Qing Liu, Huajun Yin*. Differences in root exudate inputs and rhizosphere effects on soil N transformation between deciduous and evergreen trees. Plant and Soil, 2021, 458: 277–289

lXiaomin Zhu, Dongyan Liu, Huajun Yin*. Roots regulate microbial N processes to achieve an efficient NH₄⁺ supply in the rhizosphere of alpine coniferous forests. Biogeochemistry, 2021, 155: 39–57

lWei He, Yuanshuang Yuan, Ziliang Zhang, Juan Xiao, Qing Liu, Raija Laiho, Huajun Yin*. Effect of N addition on root exudation and associated microbial N transformation under Sibiraea angustata in an alpine shrubland. Plant and Soil, 2021, 460: 469–481

lQitong Wang, Yuanshuang Yuan, Ziliang Zhang, Dongyan Liu, Juan Xiao*, Huajun Yin*. Exudate components mediate soil C dynamic through different priming mechanisms in forest soils. Applied Soil Ecology, 2021, 160, 103855

lZheng Jiang, Qitong Wang, Juan Xiao*, Ziliang Zhang, Huajun Yin. Differential responses of N benefit mediated by root exudate inputs to N addition between two subalpine forests. Rhizosphere, 2021, 19, 100404

2020年

lJunxiang Ding, Deliang Kong, Ziliang Zhang, Qin Cai, Juan Xiao, Qing Liu, Huajun Yin*. Climate and soil nutrients differentially drive multidimensional fine root traits in ectomycorrhizal-dominated alpine coniferous forest. Journal of Ecology, 2020, 108: 2544–2556

lXiaomin Zhu, Ziliang Zahang, Dongyan Liu, Yongping Kou, Qian Zheng, Mei Liu, Juan Xiao, Qing Liu, Huajun Yin*. Differential impacts of nitrogen addition on rhizosphere and bulk soil carbon sequestration in an alpine shrubland. Journal of Ecology, 2020, 108: 2309–2320

lXiaomin Zhu, Mei Liu, Yongping Kou, Dongyan Liu, Qing Liu, Ziliang Zhang, Zheng Jiang, Huajun Yin*. Differential effects of N addition on the stoichiometry of microbes and extracellular enzymes in the rhizosphere and bulk soils of an alpine shrubland. Plant and Soil, 2020, 449: 285-301

lZiliang Zhang, Qitong Wang, Yuanshuang Yuan, Qing Liu, Huajun Yin*. Rejoinder to Jones et al.‘s comments on: “Plant nitrogen acquisition from inorganic and organic sources via root and mycelia pathways in ectomycorrhizal alpine forests.”Soil Biology and Biochemistry, 2020, 136: 107517

lWanji Guo, Ziliang Zhang, Qing Liu, Juan Xiao, Huajun Yin*. Seasonal variations in plant nitrogen acquisition in an ectomycorrhizal alpine forest on the eastern Tibetan Plateau, China.Plant and Soil, 2020, 459: 79-91

2019年

lZiliang Zhang, Yuanshuang Yuan, Qing Liu, Huajun Yin*. Plant nitrogen acquisition from inorganic and organic sources via root and mycelia pathways in ectomycorrhizal alpine forests. Soil Biology and Biochemistry, 2019, 136, 107517

lZiliang Zhang, Richard P. Phillips, Wenqiang Zhao, Yuanshuang Yuan, Qing Liu*, Huajun Yin*. Mycelia-derived C contributes more to nitrogen cycling than root‐derived C in ectomycorrhizal alpine forests. Functional Ecology, 2019, 33, 346-359

2018年

lYuanshuang Yuan, Wenqiang Zhao, Ziliang Zhang, Juan Xiao, Dandan Li, Qing Liu, Huajun Yin*. Impacts of oxalic acid and glucose additions on N transformation in microcosms via artificial roots. Soil Biology and Biochemistry, 2018, 121, 16-23

lZiliang Zhang, Juan Xiao, Yuanshuang Yuan, Chunzhang Zhao, Qing Liu*, Huajun Yin*. Mycelium- and root-derived C inputs differ in their impacts on soil organic C pools and decomposition in forests. Soil Biology and Biochemistry, 2018, 123, 257-265

lZiliang Zhang, Na Li, Juan Xiao, Chunzhang Zhao, Tingting Zou, Qing Liu*, Huajun Yin*. Changes in plant nitrogen acquisition strategies during the restoration of spruce plantations on the eastern Tibetan Plateau, China. Soil Biology and Biochemistry, 2018, 119, 50-58

lWenqiang Zhao, Peter B. Reich, Qiannan Yu, Ning Zhao, Chunying Yin, Chunzhang Zhao, Dandan Li, Jun Hu, Ting Li, Huajun Yin*, Qing Liu*. Shrub type dominates the vertical distribution of leaf C: N: P stoichiometry across an extensive altitudinal gradient. Biogeosciences, 2018, 15: 2033-2053

lHuajun Yin, Wenqiang Zhao, Ting Li, Xinying Cheng, Qing Liu*. Balancing straw returning and chemical fertilizers in China: Role of straw nutrient resources. Renewable and Sustainable Energy Reviews, 2018, 81, 2695-2702

2017年及以前

lYuanshuang Yuan, Wenqiang Zhao, Juan Xiao, Ziliang Zhang, Mingfeng Qiao, Qing Liu*, Huajun Yin*. Exudate components exert different influences on microbially mediated C losses in simulated rhizosphere soils of a spruce plantation. Plant and Soil, 2017, 419: 127-140

lZiliang Zhang, Yuanshuang Yuan, Wenqiang Zhao, Heliang He, Dandan Li, Wei He, Qing Liu*, Huajun Yin*. Seasonal variations in the soil amino acid pool and flux following the conversion of a natural forest to a pine plantation on the eastern Tibetan Plateau, China. Soil Biology and Biochemistry, 2017, 105: 1-11

lDong Wang, Heliang He, Qiao Gao, Wei He, Chunzhang Zhao, Huajun Yin*, Qing Liu*. Effects of short-term N addition on soil C fluxes in alpine Sibiraea angustata scrub on the eastern margin of the Qinghai-Tibetan Plateau. Agricultural and Forest Meteorology, 2017, 247: 151-158

lZhenfeng Xu, Huajun Yin, Chunzhang Zhao, Pei Xiong, Qing Liu*. Responses of soil respiration to warming vary between growing season and non-growing season in a mountain forest of southwestern China. Canadian Journal of Soil Science, 2017, 98: 70-76

lDandan Li, Hongwei Nan, Jin Liang, Xinying Cheng, ChunZhang Zhao, HuaJun Yin, ChunYing Yin, Qing Liu*. Responses of nutrient capture and fine root morphology of subalpine coniferous tree Picea asperata to nutrient heterogeneity and competition. PLOS ONE, 2017, 11: e0187496

lHuajun Yin, P. Phillips, Rubiao Liang, Zhenfeng Xu, Qing Liu. Resource stoichiometry mediates soil C loss and nutrient transformations in forest soils. Applied Soil Ecology, 2016, 108, 248-257

lZiliang Zhang, Mingfeng Qia, Dandan Li, Huajun Yin*, Qing Liu*. Do warming-induced changes in quantity and stoichiometry of root exudation promote soil N transformations via stimulation of soil nitrifiers, denitrifiers and ammonifiers? European Journal of Soil Biology, 74: 60-68

lZiliang Zhang, Mingfeng Qiao, Dandan Li, Chunzhang Zhao, Yuejiao Li, Huajun Yin*, Qing Liu*. Effects of two root-secreted phenolic compounds from a subalpine coniferous species on soil enzyme activity and microbial biomass. Chemistry and Ecology, 2015, 31(7): 636-649

lMingfeng Qiao, Ziliang Zhang, Yuejiao Li, Juan Xiao, Huajun Yin*, Bisong Yue, Qing Liu*. Experimental warming effects on root N absorption and mycorrhizal infection in a subalpine coniferous forest. Scandinavian Journal of Forest Research, 2015, 31: 347-354

lHuajun Yin, Emily Wheeler, Richard P. Phillips. Root-induced changes in nutrient cycling in forests depend on exudation rates. Soil Biology and Biochemistry, 2014, 78: 213-221

lHuajun Yin, Yufei Li, Xiao Juan, Xinyin Cheng, Zhenfeng Xu, Qing Liu. Enhanced root exudation stimulates soil nitrogen transformations in a subalpine coniferous forest under experimental warming. Global Change Biology, 2013, 19, 2158–2167.

lHuajun Yin, Juan Xiao, Yufei Li, Zhi Chen, Xinying Cheng, Chunzhang Zhao, Qing Liu. Warming effects on root morphological and physiological traits: The potential consequences on soil C dynamics as altered root exudation. Agricultural and Forest Meteorology, 2013, 180: 287-296

lHuajun Yin, Zhi Chen, Qing Liu. Effects of experimental warming on soil N transformations of two coniferous species, Eastern Tibetan Plateau, China. Soil Biology & Biochemistry, 2012, 50, 77-84

lHuajun Yin, Zhi Chen, Yunyan Wei, Qing Liu. Nitrogen transformation in the rhizospheres of two subalpine coniferous species under experimental warming. Applied Soil Ecology, 2012, 59, 60-67

lZF Xu, Huajun Yin, Pei Xiong, Chuan Wan, Qing Liu. Short-term responses of Picea asperata seedlings of different ages grown in two contrasting forest ecosystems to experimental warming. Environmental and Experimental Botany, 2012, 77: 1-11

lLiu Q, Yin HJ, Chen JS et al. Belowground responses of Picea asperata seedling to warming and nitrogen fertilization in eastern Tibetan Plateau. Ecological Research, 2011, 26: 637-648

lHuajun Yin, Ting Lai, Xinying Cheng, Xianmin Jiang, Qing Liu. Warming effect on growth and physiology of seedlings of Betula albo-sinensis and Abies faxoniana under two contrasting light conditions in subalpine coniferous forests of western Sichuan, China. Frontiers of Forestry in China, 2009, (4): 432-442

lHuajun Yin, Qing Liu. Warming effects on growth and physiology in the seedlings of the two conifers Picea asperata and Abies faxoniana under two contrasting light conditions. Ecological Research, 2008, 23 (2): 459-469

lHuajun Yin, Qing Liu, Hai He. Seed rain and soil seed bank of Picea asperata in Subalpine coniferous plantations and natural forest in western Sichuan, China. Plant Biosystems, 2007, 141 (3): 314-322